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Sarah, a struggling young woman living in Brooklyn, agrees to volunteer as an experimental subject for a pharmaceutical company called Dynamic Health Cure to be the bearer of LEXX and make money to solve her financial troubles and take care of her mother who has Alzheimer's. A nano chip is planted into her cerebral cortex and designed to take control of the body in order to con...。在充满音乐风情的城市－加音町成长的青梅竹马北条响和南野奏，虽然升上国中后的二人个性总是不合，但却因着相同的兴趣“音乐”而有了连结。某天响与奏正巧在旧歌剧院撞见时，音乐王国“大调岛”的精灵－哈咪出现了，为了不让“小调岛”的敌人继续将传说的乐谱中“幸福的旋律”变成“不幸的旋律”音符，所以找寻能够以“高音谱记号”变身为光之美少女的响与奏。以二人合而为一的正义之心，一齐找寻飞散到人间的旋律音符，和小调岛的敌人来场音符抢夺战！。